A GUIDE TO THE STUDY OF LICHENS
/ ALBERT SCHNEIDER, M. D., Ph.D. 
A guide to the study of
lichens (1904)
Author: Schneider, Albert,
1863-1928
Subject: Lichens
Publisher: Boston, Knight
and Millet
Possible copyright status:
NOT_IN_COPYRIGHT
Language: English
Call number: 004599350
Digitizing sponsor: NCSU
Libraries
Book contributor: NCSU
Libraries
Collection: americana
Scanfactors: 30
A GUIDE TO THE STUDY OF LICHENS
/ ALBERT SCHNEIDER, M. D., Ph.D. 
Professor of Botany,
Materia Medica and Pharmacognosy, 
California College of
Pharmacy, San Francisco 
With frontispiece in color 
and 
Twenty plates 
SECOND EDITION 
BOSTON 
KNIGHT AND MILLET 
1904 
Copyright, 1898 
By Albert Schneider 
All rights reserved 
CONTENTS. 
Preface 
Introduction 
PART I. 
The General Consideration
of Lichens. 
SECTION I. 
The History of Lichenology
1 
I. From the Earliest Time
to Wallroth and 
Meyer (1825) 2 
II. From Wallroth and Meyer
(1825) to the Close 
of 1896 8 
SECTION II. 
The Uses of Lichens 16 
I. The Function of Lichens
in Nature ... 17 
II. The Economic Value of
Lichens .... 18 
SECTION III. 
What are Lichens? 25 
I. The Origin of a Lichen
29 
II. The Relation of the
Lichens to Algjs and 
Fungi 33 
in. Lichens as
Morphological Units .... 36 
VI CONTENTS. 
SECTION IV. 
The Morphology and
Physiology of Lichens ... 39 
I. The Thallus 39 
1. The Cbdstose Type 41 
2. The Foliose Type . -. 42
3. The Fbuticose Type 44 
II. The Apothecia 45 
1. The Fungal Type 46 
2. The Thalline Type 47 
III. Accessory Structures
48 
1. The Soredia 48 
2. The Cyphell-e 49 
3. The Cephalodia 50 
4. The Spermagonia 50 
SECTION V. 
The Occurrence and
Distribution of Lichens ... 52 
I. The Latitudinal and
Altitudinal Distribution 
OF Lichens 52 
SECTION VI. 
Lichens and the Naturalist
56 
I. The Collection of
Lichens 56 
II. The Study of Lichens 63
III. The Pbesebyation of
Lichens 65 
PART II. 
The Systematic Study of
Lichens. 
SECTION I. 
Systems of Classification
73 
SECTION II. 
Keys to the Study of
Lichens 76 
I. Artificial Kky to the
More Important Genera 
OCCUBBINQ IN THE UNITED
STATES ... 77 
II. Natural Key to the
Families 80 
CONTENTS. VH 
Description of Families,
Genera and Species ... 81 
I. Caliciace^ SI 
II. Cladoniace^ 87 
III. Lecideace^ 102 
IV. Gbaphidace^ . . ; 124 
V. Physciace^ 132 
VI. Parmeliace^ 144 
VII. Verrucariageje 169 
VIII. COLLEMACE^ . . 179 
IX. Pannariace^ 184 
Lichens Imperfecti, or
False Lichens 201 
The Continental Range of
the Lichens Occurring in 
THE United States 202 
Alphabetical List of the
Genera and Species Repre- 
sented in the United States
204 
General Index 225 
Plates 235 
PREFACE. 
This little work is
especially written and arranged for the 
use of amateurs in the
study of lichens. By this it is not in- 
tended to convey the idea
that the presentation of the subject- 
matter is unscientific,
incorrect or even out of date. The 
treatment of the subject is
in harmony with the most recent 
results obtained by tlie
leaders in the study of lichenology ; I 
have simply endeavored to
present these results in such a way 
that they may be
comprehended by all ; in other words, an at- 
tempt has been made to
popularize our present knowledge of 
lichens. It should,
however, be kept clearly in mind that 
only the known facts or the
completed work of science can be 
popularized. The
advance-work of science, that is, the work 
whose aim it is to make
7}ew discoveries or to correct errors, can 
never he popularized ; as
soon as this is attempted it ceases to 
be advance-work. This
statement is intended for those of the 
laity who are inclined to
speak of scientific work as " non- 
sense " or as " a
morbid taste for using big words." It is also 
intended for the serious
consideration of those would-be scien- 
tists who are too anxious
to "popularize science." It is 
further hoped that this
statement will avoid possible erro- 
neous conclusions as to the
intended purpose of this book. 
The existing
nomenclature-difiiculty does not concern or 
interest the average
student of nature. In fact, the leading 
scientific specialists are
themselves as yet wholly at sea as to 
when and where the
controversy will end. For this reason 
the citation of authorities
is omitted. Tlie names given are 
well authenticated, so that
those who have the desire and the 
X PREFACE. 
opportunity may enter into
the consideration of the synonymy 
and tlie
nomenclature-controversy. 
The lichens described are.
the more common forms occur- 
ring in the United States,
those with which tlie collector is 
likely to come in contact.
At the close is given a fairly 
complete list of the
lichens found in the United States ; this 
will prove helpful to those
who wish to make exchanges. 
The artificial key is
especially intended for the use of those 
who are not in possession
of a compound microscope. 
I take this opportunity to
express my grateful obligations 
to Dr. N. L. Britton and
Prof. Lucian M. Underwood, of 
Columbia University, who
placed at my disposal the Univer- 
sity collection of lichens.
I am also greatly indebted to my 
wife, who has kindly given
aid in correcting the manuscript 
and in reading the proof. 
Albert Schneider. 
Chicago, January, 1898. 
PREFACE TO SECOND EDITION. 
The changes to the second
edition are the addition of six 
full-page half-tone
illustrations of typical representatives of 
the three gi'eat
subdivisions of the lichens ; namely, the 
crustose, foliose, and
fruticose. These veill be of great 
value to beginners in the
study of this remai'kable group of 
plants. 
Since the issue of the
first edition no important discov- 
eries have been made in
lichenology, hence no changes have 
become necessary in the
text. 
I take this opportunity to
express my obligations to Miss 
Alice Eastwood, of the
California Academy of Science, and 
K. S. Gray, Attorney, San
Francisco, for the loan of mate- 
rial, as well as for
assistance in collecting specimens. 
Albert Schneider. 
San Francisco, August,
1903. 
INTRODUCTION. 
Lichens form a group of
plants which has been 
not so much overlooked and
neglected as misunder- 
stood and abused. The term
lichen is rather un- 
familiar, but not so the
term " moss," by which the 
plants here referred to are
quite generally known 
to layman as well as to
poet. And, indeed, the term 
moss is justifiable if we
trace it to its Scandinavian 
origin. All comparatively
small thalloid cryptogams 
— that is, flattened,
stemless and flowerless plants — 
were known as moss (most,
mossa, moos, mus). The 
term, therefore, included
lichens, liverworts and mosses 
proper. But from the
present standpoint of science 
lichens are not mosses, as
will be made clear later ; 
lichens are an independent
group of plants having no 
genetic relationship to
mosses. 
Nor must it be supposed for
a moment that lichens 
are uninteresting and
insignificant. With the one ex- 
ception of marine algae, no
plants present such truly 
beautiful and artistic
features to the amateur in nature- 
study. No group of plants
is so easily obtainable or 
Xll INTRODUCTION. 
lends itself so readily to
the critical examination of the 
student and to the
decorative fancies of the artisan. 
Neither is it necessary to
enter a plea for the kindly 
consideration of these
plants because they have been 
neglected and abused.
Lichens do not require pity : 
they are more than
competent to hold their own in 
the great struggle for
existence. Indeed, many of the 
much petted and much praised
higher plants owe their 
very existence to the
lichens. It is true, lichens can- 
not boast of an exalted
origin or noble ancestry, but 
they have gradually
advanced in a beneficent life-work, 
so that at present their
lowly origin is wholly lost 
sight of. 
Let us, therefore, obtain a
better insight into these 
plants, so that we may
judge them more fairly and as- 
sign them to their proper
position in the world of life, 
and duly credit them with
the grand work they are 
performing. Let us hope that
no one, on seeing a 
lichen, will say, " Oh
! that is a moss," or, " It's noth- 
ing ; it just grows on
trees." A lichen is as deserving 
of recognition as the oak
upon which it grows or the 
reindeer whose life it
sustains.
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